Abstract for Emilio Salinas

Gain modulation is a change in the response amplitude of a neuron that is independent of its selectivity or receptive field characteristics.  Populations of  gain-modulated neurons integrate sensory information with other kinds of sensory inputs or with proprioceptive, efference-copy, or attentional signals.  Typically this combination is multiplicative, but the key property is that it is nonlinear.  Based on theoretical results and computer simulations, it will be argued that these distributed, multi-modal representations are ideally configured to facilitate certain kinds of computations, most prominently, coordinate transformations.  Two examples will be discussed.  One from parietal cortex, where gain modulation depends on eye position and is crucial for reaching movements, and another from cortical area V4, where gain modulation depends on attention and may underlie our capacity to recognize objects independently of eye position.  It is suggested that gain modulation is a prime example of a general neural design serving a computational purpose.